There are varying stories about that. Interviews with some pretty reliable sources inside Chrysler from that time don't always agree with each other (memory is funny, and none of those guys KNEW that they were making history at the time!) But a common story is that the circle-track
426 was always intended to have a single-4 in the "bathtub" intake manifold, and that the drag version (which came along later) got the dual-quads. Since most "street hemis" went to the dragstrip and not to an oval, it made sense to sell the "street Hemi" in dual-quad form like its drag racing Race Hemi brother.